Clonal-aggregative multicellularity tuned by salinity in a choanoflagellate

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For implementers, the locking model adds a fair amount of non-trivial internal bookkeeping. Every operation must check lock state, readers must be tracked, and the interplay between locks, cancellation, and error states creates a matrix of edge cases that must all be handled correctly.

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Border Points: Each cluster has a limited number of defined "border points" – these are the gateways in and out of the cluster.